A 141-year-old Galápagos tortoise named Gramma, who lived through two world wars and 20+ U.S. presidents, has died at the San Diego Zoo.

A 141-year-old Galápagos tortoise named Gramma, estimated to have been born in 1884, has died at the San Diego Zoo. She lived through two world wars, over 20 U.S. presidents, and two global pandemics, becoming a beloved ambassador for reptile conservation. Arriving from the Bronx Zoo around 1928, she was known for her calm nature and love of cactus fruit and romaine lettuce. Officials attributed her death to age-related bone conditions. Gramma’s remarkable lifespan, potentially exceeding 140 years, is linked to her slow metabolism and biological adaptations that help resist aging. She symbolized longevity and resilience, inspiring millions worldwide.